1. Santos R., Poulet N., Cattanéo F., Besnard A. Impacts of hydro-morphological pressures on declining dace populations (Leuciscus spp.). In prep.

  2. Labeille M., Besnard A., Lefrançois E., Olivier J-M., Teichert N., Santos R., Devaux A., Bony S. Evaluation of the amphidromous fish Sicydium sp. sedentarity for biomonitoring and environmental risk assessment in the French West Indies. In prep.

  3. Santos R., Astruc G., Poulet N., Besnard A. Spatially structured freshwater fish population dynamics at the river basin district scale: implication to freshwater fish conservation in Europe. Submitted to Envionmental Sciences Europe. 

  4. Santos R., Besnard A., Bony S., Sanchez W., Devaux A. Are genotoxicants a major threat to aquatic species? Effect of paternal exposure on the level of DNA damage in sperm and  reproductive success in wild fish populations. Submitted to Ecotoxicology & Environmental Safety. 

  5. Santos R., Poulet N., Besnard A. Life-history traits correlate with freshwater fish population dynamics in common European species. Freshwater Biology, In press.

  6. Zitouni N., et al. Uptake, tissue distribution and toxicological effects of environmental microplastics in early fish juveniles (Dicentrarchus labrax). Journal of Hazardous materials, In press.

  7. Molbert A., Alliot F., Médoc V., Biard C., Meylan S., Jacquin L., Santos R., Goutte A., 2020.  Are acanthocephalans conditionally helpful parasites for chubs in polluted environments ? Environmental Science and Technology, 54 (9): 5540-5549.

  8. Goutte A., Alliot F., Budzinski H., Simonnet-Laprade C., Santos R., Lachaux V., Maciejewski K., Le Menach K., Labadie P., 2020. Trophic transfer of micropollutants and their metabolites in an urban river food web. Environmental Science and Technology, 54 (13): 8043-8050.

  9. Petitjean Q., Jean S., Côte J., Lamarins A., Lefranc M., Santos R., Perrault A., Laffaille P., Jacquin L., 2020. Combined effects of temperature increase and immune challenge in two wild gudgeon populations. Fish Physiology and Biochemistry, 46: 157-176.  doi:10.1007/s10695-019-00706-6.

  10. Simmonet-Laprade C., Budzinski H., Maciejewski K., Le Menach K., Santos R., Alliot F., Goutte A., Labadie P., 2019. Contribution of targeted and unknown precursors to the apparent biomagnification of perfluoroalkyl acids (PFAAs) in the food web of urban river. Environmental Science and Technology, doi:10.1039/C9EM00322C.

  11. Molbert N., Alliot F., Santos R., Chevreuil M., Mouchel J-M. and Goutte A., 2019. Multi-residue methods for the determination of organic micropollutants and their metabolites in fish tissues. Environmental Toxicology & Chemistry, 1218 (38): 6799-6811.

  12. Oudi A., Chokri M.A., Badraoui R., Chaabane R., Besnard A., Santos R., 2019. A multi-biomarker approach of oxidative stress reveals strong physiological impacts of pollution in Sterna hirundo in Tunisia. Marine Pollution Bulletin, 142: 196-205.

  13. Santos R, Joyeux A., Besnard A., Blanchard C., Halkett C., Bony S., Sanchez W. and Devaux A, 2017. An integrative approach to asses ecological risks of river contamination for fish populations. Environmental pollution, 220: 588-596.

  14. Santos R., Joyeux A., Palluel O., Palos-Ladeiro M., Besnard A., Blanchard C., Porcher JM., Bony S., Devaux A. and Sanchez W, 2016. Characterization of a genotoxicity biomarker in three-spined stickleback (Gasterosteus aculeatus, L.):  biotic variability and integration in a battery of biomarkers. Environmental Toxicology, 31(4): 415-426.

  15. Santos R., Palos-Ladeiro M., Besnard A., Porcher JM., Bony S., Devaux A. and Sanchez W, 2014. Kinetics response of a genotoxicity biomarker in the three-spined stickleback and implication for environmental monitoring. Ecotox. & Environ. Safety, 102: 6-11.

  16. Santos R., Palos-Ladeiro M., Besnard A., Reggio G., Vulliet E., Porcher JM., Bony S., Sanchez W. and Devaux A, 2013. Parental exposure to Methyl Methane Sulfonate of three-spined stickleback: contribution of DNA damage in male and female germ cells to further development impairment in progeny. Ecotoxicology 22(5):815-824.

  17. Santos R., Palos-Ladeiro M., Besnard A., Porcher JM., Bony S., Sanchez W. and Devaux A., 2013. Relationship between DNA damage in sperm after ex vivo exposure and abnormal embryo development in the progeny of the three spined stickleback. Reproductive Toxicology, 36:6-11.

  18. Cucchi P., Sucré E., Santos R., Leclère J., Charmantier G. and Castille R., 2012. Embryonic development of the sea bass Dicentrarchus labrax. Helgoland Marine Research, 66: 199-209.

  19. Santos R., Radford C.A., Poupin J., Brié C., Mills S.C., Galzin R. and Lecchini D., 2011. Temporal patterns in the post-larval supply of two crustacean taxa in Rangiroa atoll, FrenchPolynesia. Fisheries Science, 78 : 75-80.

  20. Wessel N., Santos R., Menard D., LeMenach K., Buchet V., Lebayon N., Loizeau V ., Burgeot T., Budzinski H., Akcha F., 2010. Relationship between PAH biotransformation as measured by biliary metabolites and EROD activity, and genotoxicity in juveniles sole (Solea solea). Marine Environmental Research, 69: S71-S73.

  21. Frederich B., Mills S.C. Denoël M., Parmentier E., Brie C., Santos R., Waqalevu V.P. and Lecchini D., 2010. Color differentiation in a coral reef fish throughout ontogeny: habitat background and flexibility. Aquatic Biology, 9 (3): 271-277.


General-public articles about my work


Actu Environnement (Please click on the logo to read the article)


Sciences et Avenir


Le Dauphiné Libéré

Peer-reviewed Publications

- Albert Einstein -

“In theory, theory and practice are the same. In practice, they are not”


  • RG

© 2015 by Raphael Santos

  • RG